PCIe x4 into x16, but what about the transfer speeds?

pcie x4 in x16 slot
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I explain in this video what is PCIe x16 on the motherboard and its uses I also explain what

So your adaptec x4 card will run at x4 in a x16 slot It's a good sign the x1 card works, but still isnt proof the x4 will work Ideally for

pcie 40 in pcie 30 slot A GTX 780R9 290 for example may suffer a performance hit but an R7 260GTX 750 may not The PCIe slots are rated as they are because of

pcie 8x slot B550M DS3H ACAMD B550 Ultra Durable Motherboard with Pure Digital VRM Solution, PCIe x16 XPG GAMMIX S50 CORE PCIe Gen4 x4 2280 Solid State Drive –